updated by 12th April 2023


[58] S. C. Qiao, I. C. Prentice, S. P. Harrison and H. Wang*: Optimality-based modelling of wheat sowing dates globally. Agricultural Systems. 206: 103608

[57] H. Wang*,  I. C. Prentice, I. J. Wright, D.I. Warton, S.C. Qiao, X.T. Xu, J. Zhou, K. Kikuzawa, N.C. Stenseth, Leaf economics fundamentals explained by optimality principles, Science Advances, 9: eadd5667

[56] Z.B. Yan, J. Sardans, J. Peñuelas, M. Detto, N.G. Smith, H. Wang, L.L. Guo, A.C. Hughes, Z.F. Guo, Calvin K. F. Lee, L.L. Liu, J. Wu,Global patterns and drivers of leaf photosynthetic capacity: The relative importance of environmental factors and evolutionary history, Global Ecology and Biogeography, 10.1111/geb.13660

[55] M.J. Zou, K. Yang, H. Liu, Y.H. Ren, J. Sun, H. Wang, S. Tan, L. Zhao, Integrating eco-evolutionary optimality principle and land processes for evapotranspiration estimation, Journal of Hydrology, 616:128855

[54] Z.Q. Zhu, H. Wang*, S.P. Harrison, I.C. Prentice, S.C. Qiao, S. Tan, Optimality principles explaining divergent responses of alpine vegetation to environmental change, Global Change Biology, 29:126-142


[53] H. Wang, S.P. Harrison, Li, M.Li, I. C. Prentice, S.C. Qiao, R.X. Wang, H.Y. Xu, G. Mengoli, G, Y.K. Peng, Y.Z. Yang, Z.B. Yan, J. Sardans, J. Peñuelas, M. Detto, N.G. Smith, H. Wang, L.L. Guo, A.C. Hughes, Z.F. Guo, Calvin K. F. Lee, L.L. Liu, J. Wu, The China plant trait database version 2, Scientific Data, 9:769

[52] H. Hunag, Z.Q. Wang, H. Wang, J. Peñuelas, J. Sardans, Ü. Niinemets, K. J. Niklas, Y. Li, J.B. Xie, and I. J. Wright, Leaf water content contributes to global leaf trait relationships, Nature Communication, 13:5525

[51] Z.Q. Zhu, H. Wang, S. P. Harrison, I.C. Prentice, S.C. Qiao, S. Tan, Optimality principles explaining divergent responses of alpine vegetation to environmental change, Global Change Biology, 29: 126-142

[50] J.X. Jin, T. Yan, H. Wang, X.L. Ma, M.Z. He, Y. Wang, W.F. Wang, F.S. Guo, Y.L. Cai, Q.A Zhu and J. Wu, Improved modeling of canopy transpiration for temperate forests by incorporating a LAI-based dynamic parametrization scheme of stomatal slope, Agricultural and Forest Meteorology, 326: 109157

[49] N. Dong, I. C. Prentice, I. J. Wright, H. Wang, O. K. Atkin, K. J. Bloomfield, T. F. Domingues, S. M. Gleason, V. Maire, Y. Onada, H. Poorter, N. G. Smith, Leaf nitrogen from the perspective of optimal plant function, Journal of Ecology, 00: 1-18

[48] H. Wang*, R.X. Wang, S. P. Harrison and I. C. Prentice: Leaf morphological traits as adaptations to multiple climate gradients. Journal of Ecology, 00: 1-12

[47] N. Dong, I. J. Wright, J. M. Chen, X. Luo, H. Wang, T. F. Keenan, N. G. Smith, I. C. Prentice: Rising CO2 and warming reduce global canopy demand for nitrogen. New Phytologist, 235: 1692-1700

[46] J. M. Chen, R. Wang, Y. H. Liu, L. M. He, H. Croft, X. Z. Luo, H. Wang, N. G. Smith, T. F. Keenan, I. C. Prentice, Y. G, Zhang, W. M. Ju, N Dong. Global Datasets of Leaf Photosynthetic Capacity for Ecological and Earth System Research. Earth System Science Data

[45] 谭深, 王焓*:基于最优性原理的普适性碳水通量耦合估算方法研究. 生态学报, 已接收


[44] S. C. Qiao, H. Wang*, I. C. Prentice and S. P. Harrison: Optimality-based modelling of climate impacts on global potential wheat yield. Environmental Research Letters, 16:114013

[43] S. Tan, X. Zhang, H. Wang, L. Yu, Y. L. Du; J. J. Yin; B. F. Wu: A CNN-based self-supervised C-band synthetic aperture radar image denoising approach. Transactions on Geoscience and Remote Sensing, accepted

[42] S. Tan, H. Wang*, I.C. Prentice, K. Yang: Land-surface evapotranspiration derived from a first-principles primary production model. Environmental Research Letters, 16:104047

[41] X. Z. Luo, T. F. Keenan, J. M. Chen, H. Croft, I. C. Prentice, N. G. Smith, A. P. Walker, H. Wang, A. R. Wang, C. G. Xu, Y. Zhang: Global variation in the fraction of leaf nitrogen allocated to photosynthesis. Nature Communication, 12:4866

[40] H. Y. Xu, H. Wang*, I. C. Prentice, S. P. Harrison, I. Wright: Coordination of plant hydraulic and photosynthetic traits: confronting optimality theory with field measurements. New Phytologist, 232: 1286–1296

[39] Z. B. Yan, Z. F. Guo, S. Serbin, G. Q. Song, Y. Y. Zhao, Y. Chen, S. B. Wu, J. Wang, X. Wang, J. Li, B. Wang, Y. T. Wu, Y. J. Su, H. Wang, A. Rogers, L. L. Liu, J. Wu: Spectroscopy outperforms leaf trait relationships for predicting photosynthetic capacity across different forest types. New Phytologist, 232: 134–147

[38] S. P. Harrison, W. Cramer, Franklin, Oskar; I. C. Prentice, H. Wang, Åke Brännström, H. de Boer, U. Dieckmann, J. Joshi, T. Keenan, A. Lavergne, S. Manzoni, G. Mengoli, C. Morfopoulos, J. Peñuelas, S. Pietsch, K. Rebel, Y. Ryu, N. Smith, B. Stocker, I. Wright: Eco-evolutionary optimality as a means to improve vegetation and land-surface models. New Phytologist, 231: 2125–2141

[37] Y. H. Ren, K. Yang, H. Wang, L. Zhao, Y. Y. Chen, X. Zhou, Z. La: The South Asia Monsoon Break Promotes Grass Growth on the Tibetan Plateau. Journal of Geophysical Research: Biogeosciences. 126: e2020JG005951

[36] H. Y. Xu, H. Wang, I. C. Prentice, S. P. Harrison, G. X. Wang, X. Y. Sun: Predictability of leaf traits with climate and elevation: a case study in Gongga Mountain, China. Tree Physiology, 41: 1336–1352


[35] C. Y. Jiang, Y. Rye, H. Wang and T. F. Keenan: An optimality-based model explains seasonal variation in C3 plant photosynthetic capacity. Global Change Biology. 26:6493–6510

[34] J. Paillassa, I. J. Wright, I. C. Prentice, S. Pepin, N. G. Smith, G. Ethier, A. C. Westerband, L. J. Lamarque, H. Wang, W. K. Cornwell and M. Vincent: When and where soil is important to modify the carbon and water economy of leaves. New Phytologist.  228: 121–135

[33] O. Franklin, S. P. Harrison, R. Dewar, C. E. Farrior, A. Brännström, U. Dieckmann, S. Pietsch, D. Falster, W. Cramer, M. Loreau, H.Wang, A. Mäkelä, K. T. Rebel, E. Meron, S. J. Schymanski, E. Rovenskaya, B. D. Stocker, S. Zaehle, S. Manzoni, M. van Oijen, I. J. Wright, P. Ciais, P. M. van Bodegom, J. Peñuelas, F. Hofhansl, C. Terrer, N. A. Soudzilovskaia, G. Midgley and I. C. Prentice: Organizing principles for vegetation dynamics. Nature Plants. 6: 444-453.

[32] S. C. Qiao, H. Wang, I. C. Prentice and P. H. Sandy: Extending a first-principles primary production model to predict wheat yields. Agricultural and Forest Meteorology. 287: 107932.

[31] H. Wang, O. K. Atkin, T. F. Keenan, N. G. Smith, I. J. Wright, K. J. Bloomfield, J. Kattge, P. B. Reich and I. C. Prentice: Acclimation of leaf respiration consistent with optimal photosynthetic capacity. Global Change Biology. 00: 1– 11.

[30] J. Kattge, et al. (724 co-authors including H. Wang): TRY plant trait database – enhanced coverage and open access. Global Change Biology. 26: 119-188.

[29] B.D. Stocker, H. Wang, N. G. Smith, S. P. Harrison, F. Keenan, D. Sandoval, T Davis and I. C. Prentice: P-model v1.0: An optimality-based light use efficiency model for simulating ecosystem gross primary production. Geoscientific Model Development. 13: 1545-1581.


[28] Y. W. Liu, S. L. Piao, T. Gasser, P. Ciais, H. Yang, H. Wang, T. F. Keenan, M. T. Huang, S. Q. Wan, J. Song, K. Wang, I. A. Janssens, J. Peñuelas, C. Huntingford, X. H. Wang, M. A. Arain, Y. Y. Fang, J. B. Fisher, M. Y. Huang, D. N. Huntzinger, A. Ito, A. K. Jain, J. F. Mao, A. M. Michalak, C. H. Peng, B. Poulter, C. Schwalm, X. T. Shi, H. Q. Tian, Y. X. Wei, N. Zeng, Q. Zhu and T. Wang: Field-experiment constraints on the enhancement of the terrestrial carbon sink by CO2 fertilization. Nature Geoscience. 12: 809-814.

[27] Y. R. Guo, C. H. Peng, Q. Zhu, M. Wang, H. Wang, S. H. Peng and H. L. He: Modelling the impacts of climate and land use changes on soil water erosion: T Model applications, limitations and future challenges. Journal of Environmental Management. 250: 109403.

[26] J. X. Jin, X. H. Ma, H. Chen, H. Wang, X. M. Kang, X. F. Wang, Y. Wang, B. Yong, F. S. Guo: Grassland production in response to changes in biological metrics over the Tibetan Plateau. Science of the Total Environment. 666: 641-651

[25] N. G. Smith, T. F. Keenan, I. C. Prentice, H.Wang, I. J. Wright, U. Niinemets, K. Y. Crous, T. F. Domingues, R. Guerrieri, F. Y. Ishida, J. Kattge, E. L. Kruger, V. Maire, A. Rogers, S. P. Serbin, L. Tarvainen, H. F. Togashi, P. A. Townsend, M. Wang, L. K. Weerasinghe, S. X. Zhou: Global photosynthetic capacity is optimized to the environment. Ecology Letters. 22: 506–517


[24] B. Yang, C. H. Peng, S. P. Harrison, H. Wei, H. Wang, Q. Zhu, M. Wang: Allocation Mechanisms of Non-Structural Carbohydrates of Robinia pseudoacacia L. Seedlings in Response to Drought and Waterlogging. Forests. 9:754

[23] H. F. Togashi, O. K. Atkin, K. J. Bloomfield, M. Bradforde, K. F. Cao, N. Dong, B. J. Evans, Z. X. Fan, S. P. Harrison, Z. Hua, M. J. Liddell, J. Lloyd, J. Ni, H. Wang, L. K. Weerasinghe, I. C. Prentice: Functional trait variation related to gap dynamics in tropical moist forests: A vegetation modelling perspective. Perspectives in Plant Ecology, Evolution and Systematics. 35: 52-64

[22] Y. Yang, H.Wang, S. P. Harrison, I. C. Prentice, I. J. Wright, C. H. Peng, G. H. Lin: Quantifying leaf trait covariation and its controls across climates and biomes. New Phytologist , 2019, 221(1): 155-168. doi: 10.1111/nph.15422

[21] H . Wang, S. P. Harrison, I. C. Prentice, Y. Z. Yang, F. Bai, H. F. Togashi, M. Wang, S. X. Zhou, J. Ni: The China Plant Trait Database: towards a comprehensive regional compilation of functional traits for land plants. Ecology. 99: 500


[20] H. Wang, I. C. Prentice, W. Cornwell, T. Davis, T. F. Keenan, I. J. Wright, C. H. Peng: A universal model for carbon dioxide uptake by plants. Nature Plants. 3: 734-741

[19] I. J. Wright, N. Dong, V. Maire, I. C. Prentice, M. Westoby, S. Díaz, R. V. Gallagher, B. F. Jacobs, R. Kooyman, E. A. Law, M. R. Leishman, U. Niinemets, R. B. Reich, L. Sack, R. Villar, H.Wang, P. Wilf: Global climatic drivers of leaf size. Science. 357: 917-921

[18] Q. A. Zhu, C. H. Peng, P. Ciais, H. Jiang, J. X. Liu, P. Bousquet, S. Q. Li, X. Q. Fang, X. L. Zhou, H. Chen, S. R. Liu, G. H. Lin, P. Gong, M. Wang, H. Wang, W. H. Xiang, J. Chen: Inter-annual Variation in Methane Emissions from Tropical Wetlands Triggered by Repeated El Niño Southern Oscillation. Global Change Biology. doi: 10.1111/gcb.13726

[17] A. Nakamura, R. L. Kitching, M. Cao, T. J. Creedy, T. M. Fayle, M. Freiberg, C. N. Hewitt, T. Itioka, L. P. Koh, K. Ma, Y. Malhi, A. Mitchell, V. Novotny, C. M. P. Ozanne, L. Song, H.Wang and L. A. Ashton: Forests and Their Canopies: Achievements and Horizons in Canopy Science. Trends in Ecology & Evolution. 32:438-451

[16] H. Wang, I. C. Prentice, T. F. Keenan, T. Davis, I. J. Wright, C. H. Peng: Photosynthetic responses to altitude: an explanation based on optimality principles. New Phytologist. 213: 976-982


[15] T. F. Keenan, I. C. Prentice, J. G. Canadell, C. A. Williams, H. Wang, M. Raupach, G. J. Collatz: Recent pause in the growth rate of atmospheric CO2 due to enhanced terrestrial carbon uptake. Nature Communication. 7: 13428 doi:10.1038/ncomms13428

[14] Q. Zhu, C. Peng, J. Liu, H. Jiang, X. Fang, H. Chen, Z. Niu, P. Gong, G. Lin, M. Wang, H. Wang, Y. Yang, J. Chang, Y. Ge, W. Xiang, X. Deng and J. S. He: Climate-driven increase of natural wetland methane emissions offset by human-induced wetland reduction in China over the past three decades. Scientific Reports. 6:38020, DOI: 10.1038/srep38020

[13] T. W. Davis, I. C. Prentice, B. D. Stocker, R. J. Whitley, H.Wang, B. J. Evans, A. V. Gallego-Sala, M. T. Sykes, W. Cramer: Simple Process-Led Algorithms for Simulating Habitats (SPLASH v.1.0): Robust Indices of Radiation, Evapotranspiration and Plant-Available Moisture. Geoscience Model Development. 10: 689-708


[12] Y. Z. Yang, Q. A. Zhu, C. H. Peng, H. Wang, W. Xue, G. H. Lin, Z. M. Wen, J. Chang, M. Wang, G. B. Liu, S. Q. Li: A novel approach for modelling vegetation distributions and analysing vegetation sensitivity through trait-climate relationships in China. Scientific Reports. doi: 10.1038/srep24110.

[11] Y. Lin, E. M. Belinda, R. A. Duursma, I. C. Prentice, H.Wang, S. Baig, D. Eamus, V. R. de Dios, P. Mitchell, D. S. Ellsworth, M. O. de Beeck, G. Wallin, J. Uddling, L. Tarvainen, M-L. Linderson, L. A. Cernusak, J. B. Nippert., T. W. Ocheltree, T. Tissue David, N. K. Martin-StPaul, A. Rogers, J. M. Warren, P. D. Angelis, K. Hikosaka, Q. Han, Y. Onoda, T. E. Gimeno, C. V. M. Barton, J. Bennie, D. Bonal, A. Bosc, M. Löw, C. Macinins-Ng, A. Rey, L. Rowland, S. A. Setterfield, S. Tausz-Posch, J. Zaragoza-Castells, M. S. J. Broadmeadow, J. E. Drake, M. Freeman, O. Ghannoum, L. B. Hutley, J. W. Kelly, K. Kikuzawa, P. Kolari, K. Koyama, J-M. Limousin, P. Meir, A. C. L. da Costa, T. N. Mikkelsen, N. Salinas, W. Sun and L. Wingate: Optimal stomatal behaviour around the world. Nature Climate Change. 5: 495-464.

[10] Y. Yang, Q. Zhu, C. Peng, H. Wang, H. Chen: From plant functional types to plant functional traits: A new paradigm in modelling global vegetation dynamics. Progress in Physical Geography. 39: 514-535.

[9] T-T. Meng, H.Wang, S. P. Harrison, I. C. Prentice, J. Ni and G. Wang: Responses of leaf traits to climatic gradients: adaptive variation vs. compositional shifts. Biogeosciences. 12: 1-14.


[8] H.Wang, I. C. Prentice, T. W. Davis: Biophysical constraints on gross primary production by the terrestrial biosphere. Biogeosciences. 11: 5987-6001.

[7] B. J. Atwell, H. Wang, A. P. Scafaro: Could abiotic stress tolerance in wild relatives of rice be used to improve Oryza sativa? Plant Science. 215-216: 48-58.


[6] H. Wang: A multi-model assessment of climate change impacts on the distribution and productivity of ecosystems in China. Regional Environmental Change. 14: 133-144.

[5] H. Wang, I. C. Prentice, J. Ni: Data-based modelling and environmental sensitivity of vegetation in China. Biogeosciences. 10: 5817-5830.


[4] H. Wang, I. C. Prentice, J. Ni: Global change and primary production in China: contrasting light- and water-use efficiency models. Biogeosciences. 9: 4689-4705.

[3] D. I. Kelley, I. C. Prentice, S. Harrison, H.Wang, M. Simard, J. B. Fisher, K. Willis: A comprehensive benchmarking system for evaluating global vegetation models. Biogeosciences. 10: 3313–3340.


[2] H.Wang, J. Ni, I. C. Prentice: Sensitivity of potential natural vegetation in China to projected changes in temperature, precipitation and atmospheric CO2. Regional Environmental Change. 11: 715-727.

[1] I. C. Prentice, T. Meng, H. Wang, S. Harrison, J. Ni, G. H. Wang: Evidence for a universal scaling relationship of leaf CO2 drawdown along an aridity gradient. New Phytologist. 190: 169-180.